Heterostegina papyracea Seguenza, 1880 from the upper Miocene of Cessaniti (Vibo Valentia, Calabria, southern Italy)

Several specimens of Heterostegina papyracea Seguenza were collected near Cessaniti (Vibo Valentia) in upper Tortonian-lower Messinian sandstones with abundant Clypeaster and small pectinids.The biometric analysis of the inner features of the specimens of H. papyracea leads us to insert it into the H. complanata lineage. In the Mediterranean area H. papyracea probably represents the last species of this group. The H. costata lineage is represented in Spain by H. gomezangulensis Perconig.